'Queen' Gramma the Tortoise, the Oldest Animal at the San Diego Zoo, Dies at the Estimated Age of 141

Gramma the Galápagos tortoise died on Nov. 20, 2025. The oldest animal at the San Diego Zoo was believed to be around 141 ...

Gramma, a Galápagos Tortoise Older Than the San Diego Zoo, Dies at About 141

Gramma, who died at the zoo on Thursday, was believed to have been born in 1884, when Chester A. Arthur was in the White ...

Gramma the Galápagos tortoise, San Diego Zoo's oldest resident, dies at about 141

Beloved Galápagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died at about 141, the zoo says, adding that ...

Oldest resident of San Diego Zoo, Gramma the Galapagos tortoise, dies at estimated 141 years old

After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise ...

Gramma the Galápagos tortoise, an original San Diego Zoo resident, dies at 141

Gramma the tortoise, believed to be 141 years old and the oldest resident of the San Diego Zoo, was euthanized after suffering from progressive bone deterioration because of her age.

Gramma the tortoise, oldest animal at the San Diego Zoo, dies

Gramma, a Galapagos tortoise that was the oldest living animal at the San Diego Zoo, has died at an estimated 141 years of ...

Gramma, San Diego Zoo’s Galápagos tortoise, dies at 141

She had “ongoing bone conditions related to advanced age” that recently progressed and the care team “made the compassionate ...